After the death of King Cretheus, the Aeolian Pelias usurped the Iolcan throne from his half-brother Aeson and became king of Iolcus in Thessaly (near the modern city of Volos). Because of this unlawful act, an oracle warned him that a descendant of Aeolus would seek revenge. Pelias put to death every prominent descendant of Aeolus he could, but spared Aeson because of the pleas of their mother Tyro. Instead, Pelias kept Aeson prisoner and forced him to renounce his inheritance. Aeson married Alcimede, who bore him a son named Jason. Pelias intended to kill the baby at once, but Alcimede summoned her kinswomen to weep over him as if he were stillborn. She faked a burial and smuggled the baby to Mount Pelion. He was raised by the centaur Chiron, the trainer of heroes.
When Jason was 20 years old, an oracle ordered him to dress as a Magnesian and head to the Iolcan court. While traveling Jason lost his sandal crossing the muddy Anavros river while helping an old woman (Hera in disguise). The goddess was angry with King Pelias for killing his stepmother Sidero after she had sought refuge in Hera's temple.
Another oracle warned Pelias to be on his guard against a man with one shoe. Pelias was presiding over a sacrifice to Poseidon with several neighboring kings in attendance. Among the crowd stood a tall youth in leopard skin with only one sandal. Pelias recognized that Jason was his nephew. He could not kill him because prominent kings of the Aeolian family were present. Instead, he asked Jason: "What would you do if an oracle announced that one of your fellow-citizens were destined to kill you?". Jason replied that he would send him to go and fetch the Golden Fleece, not knowing that Hera had put those words in his mouth.
Jason learned later that Pelias was being haunted by the ghost of Phrixus. Phrixus had fled from Orchomenus riding on a divine ram to avoid being sacrificed and took refuge in Colchis where he was later denied proper burial. According to an oracle, Iolcus would never prosper unless his ghost was taken back in a ship, together with the golden ram's fleece. This fleece now hung from a tree in the grove of the Colchian Ares, guarded night and day by a dragon that never slept. Pelias swore before Zeus that he would give up the throne at Jason's return while expecting that Jason's attempt to steal the Golden Fleece would be a fatal enterprise. However, Hera acted in Jason's favour during the perilous journey.
Jason arrived in Colchis (modern Black Sea coast of Georgia) to claim the fleece as his own. It was owned by King Aeetes of Colchis. The fleece was given to him by Phrixus. Aeetes promised to give it to Jason only if he could perform three certain tasks. Presented with the tasks, Jason became discouraged and fell into depression. However, Hera had persuaded Aphrodite to convince her son Eros to make Aeetes's daughter, Medea, fall in love with Jason. As a result, Medea aided Jason in his tasks. First, Jason had to plow a field with fire-breathing oxen, the Khalkotauroi, that he had to yoke himself. Medea provided an ointment that protected him from the oxen's flames. Then, Jason sowed the teeth of a dragon into a field. The teeth sprouted into an army of warriors (spartoi). Medea had previously warned Jason of this and told him how to defeat this foe. Before they attacked him, he threw a rock into the crowd. Unable to discover where the rock had come from, the soldiers attacked and defeated one another. His last task was to overcome the sleepless dragon which guarded the Golden Fleece. Jason sprayed the dragon with a potion, given by Medea, distilled from herbs. The dragon fell asleep, and Jason was able to seize the Golden Fleece. He then sailed away with Medea. Medea distracted her father, who chased them as they fled, by killing her brother Apsyrtus and throwing pieces of his body into the sea; Aeetes stopped to gather them. In another version, Medea lured Apsyrtus into a trap. Jason killed him, chopped off his fingers and toes, and buried the corpse. In any case, Jason and Medea escaped.
In Corinth, Jason became engaged to marry Creusa (sometimes referred to as Glauce), a daughter of the King of Corinth, to strengthen his political ties. When Medea confronted Jason about the engagement and cited all the help she had given him, he retorted that it was not she that he should thank, but Aphrodite who made Medea fall in love with him. Infuriated with Jason for breaking his vow that he would be hers forever, Medea took her revenge by presenting to Creusa a cursed dress, as a wedding gift, that stuck to her body and burned her to death as soon as she put it on. Creusa's father, Creon, burned to death with his daughter as he tried to save her. Then Medea killed the two boys that she bore to Jason, fearing that they would be murdered or enslaved as a result of their mother's actions. When Jason came to know of this, Medea was already gone; she fled to Athens in a chariot sent by her grandfather, the sun-god Helios.
Later Jason and Peleus, father of the hero Achilles, attacked and defeated Acastus, reclaiming the throne of Iolcus for himself once more. Jason's son, Thessalus, then became king.
As a result of breaking his vow to love Medea forever, Jason lost his favor with Hera and died lonely and unhappy. He was asleep under the stern of the rotting Argo when it fell on him, killing him instantly.
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